Gary, great explanation!  
Suzi, the Polar website explains this in depth.   Here's a link: 
 
https://www.polarusa.com/targetzone/default.asp for figuring mhr (max heart rate) out.  Another good site is: 
 
https://www.bodyforlife2.com/max_heart_rate.htm 
 
Basically, you cannot rely on just the 'formula' as every body is different.  It is a good starting point but, I found, if I used it only I'd be moving way less.  I hope this helps.